We introduce Nakayama functors for coalgebras and investigate their basic properties. These functors are expressed by certain (co)ends as in the finite case discussed by Fuchs, Schaumann, and Schweigert. This observation allows us to define Nakayama functors for Frobenius tensor categories in an intrinsic way. As applications, we establish the categorical Radford S4-formula for Frobenius tensor categories and obtain some related results. These are generalizations of works of Etingof, Nikshych, and Ostrik on finite tensor categories and some known facts on co-Frobenius Hopf algebras.
